Here's not the case as TR2 act on the 2n3904/06
bases before TR3-Tr6 conveyor act on the emitters so the speed of switching is determined by the Tr3-Tr6 delay mainly and they are clearly slower than 2n3904/06. That sequence is deliberate to forcefully discharge the base of the trz that needs to shut off, the circuit doesn't rely only on the underbias for that matter .As with the capacitances involved Low cob trz are usually low on beta too because they are mainly destined for common base connections...in common base connection miller's capacitance is just a fraction of cob as the voltage gain is below 1X so cob or rather ccb in this case is irrelevant .
I remember that the
UHF-VHF amplifiers I was building in childhood to catch the russian's channels barely had 10db gain per 1 trz stage made with some BFR90...91 trz used in common emitter connection so its CCE would be important while multiplied with the stage voltage gain which was like 1/5 of the dc current gain thus 10 db gain was kinda big deal at 800Mhz .
BF422/423 would be a more natural choice than BFR trz which I'm not sure how stable can be in audio while bf422/3 definitely work in audio, but I'm pretty sure VHF trz can easily be toast due to very low CE max voltages common to these types of transistors.Unless you're going to use high power RF trz you won't get big VCE ranges...and big power trz are usually used in common base connection or cascoded "two on a die" to reduce the influence of their inherently higher cob ...