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    Building, troubleshooting and testing of these amplifiers should only be
    performed by someone who is thoroughly familiar with
    the safety precautions around high voltages.

voltages making no sense - help!

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V2 has a special case where the load impedance "downstream" of the valve's own anode resistor is so high that it can be ignored - excellent! We need only mark two points on the axes and draw a straight line. Your posted curves are close but not exactly on classic American curves for 12AX7, plenty close enough, and the missing grid voltage curves' markings are in 0.5 VDC increments from zero (left).

Your operating "curve" (although we're drawing a straight line resistive approximation, the actual loadline is slightly elliptical because stray capacitance "fattens" the line, linearly variable with frequency; not a significant issue within the audio range in this particular case, but surprisingly significant in other cases) is a straight line from a point on the X axis at B+ (let's call it +378 VDC) and a point on the Y axis at +378 VDC / 301K Ohm (the anode DC load resistor). This is the current that would occur if the valve were able to conduct completely - a dead short. So, the reverse slope of this line is 301K Ohm.

As the voltage on the grid (blue lines) varies, anode voltage (on the red loadline) varies with it. 12AX7s are amazingly linear, so some amount of grid voltage change causes pretty much the same amount of anode voltage change at various places along the loadline, but there are definitely better and best places to operate.

If this is too simplistic, or not the actual thrust of your question, please restructure it and I'll try my best.

All good fortune,
Chris
 
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Well actually on further analysis and testing the culprit was actually a 0.082uF 400V Jensen PIO I had on as a bypass I had forgotten about on the other side of the PCB to replace the original bypass cap C40. Oh well, at least I am learning something!
 
For sure.

Jensen Aluminium Foil, Paper in Oil, in an Aluminium can: These leak DC in my case about 700 mV from 170V, which upsets all of the voltages across the valve. I tried both the 400V and 630V versions

Arizona Capacitors Blue Cactus : These work and 'leak only' 6-10 mV, this gets me the correct voltages across the valve as expected.


I also tried these and they work too: Metalized Polyester Film DME Orange Drops [30v]



 
For sure.

Jensen Aluminium Foil, Paper in Oil, in an Aluminium can: These leak DC in my case about 700 mV from 170V, which upsets all of the voltages across the valve. I tried both the 400V and 630V versions

Arizona Capacitors Blue Cactus : These work and 'leak only' 6-10 mV, this gets me the correct voltages across the valve as expected.


I also tried these and they work too: Metalized Polyester Film DME Orange Drops [30v]
DME orange drops are 630V