Ketje, Agreed, nevertheless even with strangely off bias points, the circuit should work quite well (with the possible exception should the V7A plate get too low in voltage). Also looks like my sims, and your bias point calculations are in reasonable agreement. My models may not be that bad.
Just a thought, if filament voltages were low, the tubes would have lower conductance for a given gate drive. This could explain the peculiar biases. Why the filament voltage would be low for the original design would still be a mystery.
Anyway, the filament voltage should be checked with all tubes lit up.