Very specific recommendations:
[1] Pentode cathode resistor 68 ohms, change to 220 ohms.
[2] Bypass pentode plate transformer primary with 47K, 1 watt resistor in parallel with primary. Carbon composition is just fine for this purpose.
Theory behind #1: every possibility that transformer is running close to saturation. Drop amperes through it, without substantially changing gain. Should have significant effect if hypothesis is correct.
Theory behind #2: inductive load becomes high enough to emphasize transients. Providing a simple resistive "knee" in the response curve gives pentode a real non-transformative load to chew on. May well, by itself, solve problem. Suggestion above to use a 100K pot is good. Be careful - at lower ohm settings, may burn out due to excessive power dissipation. You can put a 15K resistor in series with it, to prevent, but the 15K needs to be 5 watt.
I have a third and fourth suggestion, but they're less likely, and require more difficult wiring changes on the output MOSFETs. By the way - the output mosfets at 2.1 volts may be significantly over-biased. This can cause HF clipping, as they are not particularly linear MOSFETS.
Good luck.
Hi GoatGuy, tried both your suggestions, and I hate to say this : no audible difference. So ........ what are your 3rd and 4th ideas ?? Having said which, the amp sounds extraordinary and I really appreciate your help.