Quad 33 popcorn/crackling in 1 channel

I recently got a Quad 33 for almost free, to be used with my 303.
It sounds fine, everything works, however, there’s a faint “popcorn” noise to be heard when you put your ears against the speakers. The noise is independent from volume, and inaudible when I play music.

When I change the position of the M12017 amplifier boards, the problem changes too, to the other speaker. Therefore I think I narrowed it down to one of these M12017 boards.

Firstly, I renewed C401, C405 and C406. That did not solve the problem, and I put them back.

I checked the supply rail and saw nothing strange there, I would be surprised since the issue is strictly in one channel.

Any suggestions what to check next? Any “known Quad 33” issues that sounds like mine?

Thanks!

(this question was also posted in vintage radio forum)
0D9058ED-804B-4D4C-A8AB-D304EE6310C9.jpeg
 
C400 and Tr400 would be next on my list...other than that if you have access to a scope to narrow down where the "popcorn" is coming from and substitution is the only thing you can do...try subbing all the BC 109's one at a time.