• WARNING: Tube/Valve amplifiers use potentially LETHAL HIGH VOLTAGES.
    Building, troubleshooting and testing of these amplifiers should only be
    performed by someone who is thoroughly familiar with
    the safety precautions around high voltages.

Heater reference

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If one cathode is pulled all the way down to -400 Volts, it is a reasonable assumption the other cathode does not rise above -300 volts. With the heaters biased reference at the -400 Volt rail. the voltage from cathode to heater should always fall between 0 to 100 volts.
 
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